Carl Ferraris and Mario di Pinna after an in-depth review
of the relevant literature, have published a comprehensive
list of all family-group and other suprageneric names proposed
for the order Siluriformes. Their findings will undoubtedly
cause a few groans from hobbyists, as we now have to learn
some new names. Still other hobbyists may be disappointed
that the authors chose not to retain such families as the
Ageneiosidae. However, this new paper is important. While
the rules that govern generic and specific names are very
stringent, family level names do not come under such scrutiny.
For many years it has been nearly impossible to know exactly
how many Catfish families legally (legal as defined by the
International Code of Zoological Nomenclature) exist. Hopefully,
this paper will provide some, at least temporary, stability
to the number of catfish families.
A complete discussion of taxonomic terminology is beyond
the scope of this paper, but a brief introduction to some
of the terminology is certainly in order. Phylogeny (from
the Greek phylon = tribe and genesis = origin) is the evolutionary
history of a species or group of related species. These
genealogies trace evolutionary relationships. Reconstructing
phylogenetic history is part of the scope ot systematics,
the branch of biology concerned with the diversity of life.
Systematics encompasses taxonomy, which is the identification
and classification of species. A family, as used in this
paper, is a taxonomic grouping below order and above genus.
Family names for plants always end in acae while those of
animals, like catfishes, always end in idae.
The family level is an important classification for hobbyists.
If the aquarist knows which family a fish belongs to they
can immediately make inferences about the fish's diet, adult
size, natural habitat, and other important information for
captive maintenance. This is especially true in those cases
where the aquarist is lucky enough to make when of those
truly rare finds. At these times, it can be almost impossible
to identify the fish to genus, let alone species and identification
to family is the best we can hope for.
Below is a list of the 35 catfish families accepted by the
authors after their review of the literature. I will make
a few comments under each family to point out any radical
changes or changes that affect popular aquarium catfishes.
I will also point out common family names where they exist
or have been confused and comment on the family's availability
in the aquarium trade. I have taken the liberty of suggesting
a few common names for some of the newer families.
Akysidae
Asian family rarely imported
for the aquarium trade. Collectively known as the Asian
banjo cat-fishes (Ferraris, 1991: 164) or stream catfishes
(Jayaram, 1999: 266). The family Parakysidae, (Roberts,
1989), pustulous catfishes (Kottelat et al 1993: 105),
is included in this family.
Amblycipitidae
Very rarely imported family
from Asia. Collectively known as the loach-catfishes
(Burgess, 1989: 107).
Amphiliidae
Very rarely imported family
found in Africa. Collectively known as the African hillstream
cat-fishes (Burgess: 109) or mountain catfishes (Skelton,
1993: 218).
Anchariidae
Erected by Glaw and Vences,
1994 to accomodate Ancharius Steindachner, 1881.
The family was proposed by de Pinna in an unpublished
thesis. I have not seen the original description and
thus can not add much. The family is found on Madagascar.
I am not aware of any common name or importations.
Andinichthyidae
Proposed by Gayet, 1988 to
accommodate the fossil genus and species Andinichthys
bolivianensis from South America. Obviously not
an aquarium import.
Ariidae
A world-wide family known
as the sea catfishes (Burgess, 1989: 158; Ferraris,
1991: 82) or shark catfishes (Baensch & Riehl, 1997:
434) even though some members are restricted to freshwater.
Allen (1 989:47) uses the term fork-tailed catfishes.
Known in the hobby mainly for the Arius species
imported as "shark catfish"
Aspredinidae
A South American family known
as the banjo catfishes (Burgess: 295). A number of species
are common imports.
Astroblepidae
South American hillstream
catfishes (Burgess: 446) or Andes catfishes (Ferraris:
166). Restricted to South America and likely never imported.
Auchenipteridae
Collectivly known as the driftwood
catfishes (Burgess: 226). The family Ageneiosidae, the
slopehead cat-fishes, appears to be included under this
family, but is not specifically discussed. Imports range
from the common (e.g. the Zamora or midnight catfish)
to the rare (e.g. the jaguar catfish) to the very rare.
Austroglanididae
Collectively known as the
rock catfishes (Skelton, 1993: 215), this family was
erected by Mo, 1991 to accomodate the south African
genus Austroglanis. All three species contained
in the family are rare in nature and threatened or endangered
by habitat destruction.
Bagridae
The family Bagridae, after
Mo's 1991 revision, is now an exclusively Asian family
with the exception of a single genus, Bagrus,
that occurs in Africa. The family Olyridae, bannertail
catfishes (Burgess: 153), is also now included in the
Bagridae. Many members of the family are common to rare
imports. Collectively known as the bagrid catfishes.
Callichthyidae
A South American family well
known in the aquarium hobby especially for the members
of the genus Corydoras. Collectively known as
the armored catfishes Riehl & Baensch (1991: 453).
Cetopsidae
A South American family known
as the whale catfishes (Burgess: 289). One or two species
are rarely imported. Helogenidae, the marbled catfishes
(Burgess: 287), is included in Cetopsidae.
Chacidae
A small Asian family of three
species that are uncommonly imported. Collectively known
as the frog-mouth catfishes (Burgess: 151). The common
name angler catfishes has also been applied (Ferraris:
109) but should not be used as it appears that Chaca
do not angle (Linder, 1998: 3).
Clariidae
Collectively known as the
labyrinth catfishes (Burgess: 135), walking catfishes
(Baensch & Riehl, 1997: 484), and air-breathing
catfishes (Skelton: 227) this family is widely distributed
throughout Africa and Asia. The U.S. Fish and Wildlife
Service has declared them "injurious wildlife"
and their import is banned (Ferraris: 113). Occasional
specimens come in to the U.S. as by-catch or contaminants.
Claroteidae
Erected by Mo, 1991 to accommodate
most of the African genera that were formerly of the
Bagridae. This family is composed of 13 genera and over
90 species and are referred to simply as Claroteid catfishes
(Skelton: 211). A few members of the genera Auchenoglanis,
Parauchenoglanis, Chrysichthys, Clarotes,
Gephyrogianis, Lophiobagrus, and Phyilonemus
are uncommonly to rarely imported (Glaser, 2000).
Cranoglanididae
Known as Chinese catfishes
(Burgess: 72) this Asian family contains but one species
that has not been imported.
Diplomystidae
A South American family known
as the Patagonian catfishes (Burgess: 23). This small
family, with about four species, has not been imported.
They are found in swift cool streams in Chile and Argentina.
Doradidae
A popular South American family
known as the talking catfishes (Burgess: 199 Ferraris:
114) and thorny catfishes (Riehl & Baensch, 1991:)
Importations range from common (the so called raphaels)
to rare
Erethistidae
This Asian family consists
of Conta, Laguvia, Pseudolaguvia,
Erethistoides, Hara, and Erethistes
which are genera removed from the family Sisoridae.
A few genera are uncommon to rare imports. No common
name has been applied to this family. The family was
erected by de Pinna in 1996. However, some subsequent
authors (e.g. Jayaram, 1999) have not followed de Pinna's
findings, while others have (e.g. Grant, 1999: 9). Hopefully,
Ferraris and de Pinna's 1999 paper will stabilize the
use of this family.
Heteropneustidae
This small Asian family are
referred to as airsac catfishes (Burgess, 148), fossil
catfishes, and stinging catfishes (Ferraris, 121). Imports
have become rare in recent years. Perhaps because of
the restrictions placed on the closely related family
Clariidae.
Hypsidoridae
Erected for the catfish
genus Hypsidoris Lundberg and Case, 1970.
lctaluridae
A primarily North American
family. Although maintained by a few specialist aquarists
there is no organized commercial trade in place for
the aquarium hobby. The only species commercially traded
are juvenile channel cattishes that are brought into
the hobby trade by aquaculture enterprises. Riehl and
Baensch (1991: 453) use horned pouts as a common name
and Burgess (1989: 26) uses bullhead cattishes . However,
this latter name is normally applied only to members
ot the genus Ameiurus. Members of Ictalurus
are commonly known as forktail catfishes, Pylodictus
as the flathead cattish, and the largest genus Noturus
as the madtoms. The common name should be standardized
as North American catfishes as it is the only family
native to the continent.
Loricariidae
A large South American family
that is very popular in the aquarium hobby. Common names
include armoured sucker-mouthed catfishes and armoured
catfishes (Ferraris, 126). Armour-plaited catfishes
(Riehl and Baensch 1991: 453), suckermouth catfishes
(Burgess: 368), sucker-mouthed armoured catfishes (Innes,
1966: 285), and, of course, the plecos. Some scientific
papers use armoured catfishes while lsbrücker and
Nijssen (two scientists that have done a lot of taxonomic
work on the family) consistently use the term mailed
catfishes. With so many common names in use, it is impossible
to suggest a single universal common name
Melapteruridae
A small African family known
as the electric cat-fishes (Burgess: 155, Riehl &
Baensch, 1991:453). At least one species is an uncommon
import.
Mochokidae
A large and diverse African
family. Burgess (182) uses the term upside-down catfishes
but only a very few members of a single genus swim in
this manner. Also, at least one bagrid is an upside-down
swimmer. Skelton (240) divides the family into squeakers
(Synodontis) and suckermouth catlets (Chiloglanis)
which is certainly more descriptive. Because the family
is so diverse as to defy a common descriptive name,
the term mochokid catfishes should be used.
Nematogenyidae
A small South American family
consisting of a single genus. I am unaware of any importations
of this family for the aquarium trade. The common name
should be worm catfishes which follows from the family
name and is descriptive of the family.
Pangasiidae
An Asian family referred to
as the shark cat-fishes by Burgess (100). This term
has also been applied to the Ariidae. One species (Pangasius
hypophthalmus, the irridescent shark) is common.
Other species are showing up in the trade as the result
of aquaculture programs.
Pimelodidae
A large and popular South
American family. Common names include antenna catfishes
(Burgess: 243) and flat hosed (nosed?) catfishes (Riehl
and Baensch, 1991: 453). Imports are common to rare
depending on the species with Pseudopimelodus,
Microglanis, and Pimelodus the most common.
Hypophthalmidae, lookdown catfishes (Burgess: 293),
are included in Pimelodidae.
Plotosidae
A widely distriduted family
that includes marine species. Common names include tandan
catfishes (Burgess: 171), eel-tailed catfishes (Ferraris:
157). The name eel-tailed catfishes is the most commonly
accepted (Allen, 1989: 55 Jayaram, 1999: 317, Kottelat
et al 1993: 113). Only one species, Plotosus lineatus,
is a common import for the marine hobby.
Schilbidae
A medium sized family found
in Africa and Asia. Sometimes spelled Schilbeidae (Burgess:
87 Jayaram: 249). Common names include glass catfishes
(Burgess: 87, Riehl & Baensch 1991: 453) and butter
catfishes (Skelton: 224). However, the term schilbid
catfishes is more descriptive as very few members are
transparent and the term glass catfishes is also often
applied to the Siluridae. Importation of one Eutropieiius
and one Schilbe species is common, but all
others, especially those from Asia, are rare.
Scoloplacidae
A small family found in South
America and referred to as spiny dwarf catfishes (Burgess:
450). Only very rarely imported and usually only as
by-catch.
Siluridae
A large family found from
Europe (two species) through Asia. Commonly known as
sheat cat-fishes (Burgess: 74), glass catfishes (Ferraris:
161), old world catfishes (Riehi & Baensch 1991:
453), and sheath (Baensch & Riehl, 1997: 576). Sheat
catfishes is the term preferred in most scientific works
and should be used to refer to the family. Importation
of Southeast Asian species ranges from common (various
Kryptopterus) to uncommon (e.g. Ompok)
to rare (e.g. Belodontichtys) with all other
species rarely, if ever, imported.
Sisoridae
A large Asian family commonly
known as Asian hillstream catfishes (Burgess: 119) and
sucking catfishes (Kottelat et al: 106). Asian hill
stream catfishes in the most widely used name. Imports
are generally rare. This appears to be mainly due to
the high oxygen requirements and cooler temperatures
demanded by most species. Neither of these requirements
is conducive to commercial shipping.
Trichomycteridae
A large South American family
known as parasitic catfishes (Burgess: 305). This name
is probably undeserved as most species are not parasitic.
However, it is likely to remain in usage. Imports are
rare and infrequent. These fishes undeserved reputation
scares off many would be importers.
References:
Allen G. A. 1989. Freshwater Fishes of Australia. TFH Publications;
Neptune City, New Jersey. 240 pp., 64 pls.
Baensch, H. A. & A. Riehl. 1997. Aquarium Atlas Volume
Two. Mergus; Germany: 1212 pp. Second English Edition.
Burgess W. E. 1989. An Atlas of Freshwater and Marine Catfishes.
TFH Publications; Neptune City, New Jersey. 784 pp.
Burgess W. & L. Finley. 1996. An Atalas of Freshwater
and marine Catfishes: Update. Tropical Fish Hobbyist. Vol.
XLV No. 2 (#488) October, 1996
Ferraris C. J. 1991. Catfish in the Aquarium. Tetra; Morris
Plains, New Jersey. 199 pp.
Ferraris C. J. & M. C. de Pinna. 1999. Higher Level
Names for Catfishes (Actinopterygii: Ostariophysi:
Siluriformes). Proceedings of the California Academy of
Sciences. Vol. 51 No. 1: 1-17.
Jayaram, K. C. 1999. The Freshwater Fishes of the Indian
Region. Narendra Publishing House; Delhi, India. 551 pp.,
28 pls.
Kottelat, M., A. J. Whitten, S. N. Kartikasari & S.
Wirjoatmodjo, 1993. Freshwater Fishes of Western Indonesia
and Sulawesi. Periplus Editions, Hong Kong, 221 pp., 84
p15.
Linder, R. 5. 1998. The Catfishes of Asia: Chacidae. Newsletter,
Northern Area Catfish Group. Issue No. 3: 3-6.
Mo, T. 1991. Anatomy, Relationships, and Systematics of
the Bagridae (Teleostei: Siluroidei) with a Hypothesis of
Siluroid Phylogeny. Koeltz, Koenigstein, viii +21 6pp.
Pethiyagoda R. 1991. Freshwater fishes of Sri Lanka. Wildlife
Heritage Trust of Sri Lanka, Colombo, xiv+362pp.
Riehl, R. & H. A. Baensch. 1994. Aquarium Atlas. Mergus;
Hong Kong. 992 pp. Fourth English edition, reprinted.
Riehl, R. & H. A. Baensch. 1996. Aquarium Atlas Volume
Three. Mergus; Germany. 1104 pp.
Roberts T. R. 1989. The freshwater fishes of western Borneo
(Kalimantan Barat, Indonesia). Mem. California Acad. Sci.,
14: 1-21a
Skekon, P. 1993. A Complete Guide to the Freshwater Fishes
of Southem Africa. Southern Book Publishers; Halfway House.
388 pp.